K. Lloyd Curriculum Vitae

University of Tennessee • Microbiology Department • M409 Walters Life Sciences • Knoxville, TN 37996

Ph: 865-974-4224 • Fax: 865-974-4007 • klloyd@utk.edu

_____________________________________________________________________________________

Education

University of North Carolina, Chapel Hill, M.S. May 2005, Ph.D. Aug 2009, Marine Sciences 

Swarthmore College, Swarthmore, PA. B.A. June 2000, Biochemistry

_____________________________________________________________________________________

Professional Experience

Assistant Professor, Department of Microbiology, University of Tennessee                       Feb 2012 - current

Postdoctoral Researcher, Center for Geomicrobiology, Aarhus U., Denmark              Sept 2009 – Dec 2011

Guest Investigator, Woods Hole Oceanographic Institution                                         Sept 2001 – Aug 2002                                                                                                    

Project Coordinator, Northwestern University, Dept. of Medicine                                Sept 2000 – June 2001

_____________________________________________________________________________________

Awards and Fellowships

Simons Early Career Investigator in Marine Microbial Ecology and Evolution Award (2016)

NASA Exobiology Grant, Lead PI (2016)

Sloan Fellowship, Alfred P. Sloan Foundation (2015)

NSF Dimensions of Biodiversity Grant, Co-PI (2015) 

NSF Expedition Objective Research Grant, Lead PI (2014)

Deep Carbon Observatory Census of Deep Life DNA sequencing grant, NSF and Sloan Foundation (2014)

Center for Dark Energy Biosphere Investigations Biomolecular Research Grant, Lead PI, NSF (2013-2014)

Center for Dark Energy Biosphere Investigations Small Research Grant, Lead PI, NSF (2012-2013)

Holger Jannasch Visiting Scholar Award, Woods Hole Oceanographic Institution (2012)

U. S. EPA STAR Fellowship for tuition, stipend, research materials & travel (2006-2009)

North Carolina Space Grant Fellowship for research materials (2006-2007)

NASA Astrobiology Institute Travel Fellowship for the International Workshop on Microbial Life under Extreme Energy Limitation. Aarhus, Denmark (2007)

Graduate Travel Stipend, from U. North Carolina for ISME Meeting, Vienna, Austria   (2006)

NASA Astrobiology Institute Scholarship for the NASA Graduate Winter School (2005)

Doctoral International Travel Fellowship from the University Center for International Studies to work at the Max Plank Institute for Marine Microbiology in Bremen, Germany (2005)

NASA Astrobiology Institute Scholarship for the NASA Graduate Summer School (2004)

Agouron Institute Scholarship for the International Geobiology Summer Course (2003)

University Merit Assistantship from the University of North Carolina (2002-2003)

_____________________________________________________________________________________

Publications

     Bird, J. T., B. J. Baker, M. Podar, K. G. Lloyd. Similarities between distantly related members of the Altiarchaeales suggest novel Order of Archaea is predominantly autotrophic. Submitted. 

     Buongiorno, J., S. Turner, G. Webster, M. Asai*, A. Shumaker*, T. Roy, A. Weightman, A. Schippers, K. G. Lloyd. Quantification of deep subseafloor bacteria and archaea – an interlaboratory comparison. In revisions

     Kevorkian, R. T., J. Bird, A. Shumaker, K. G. Lloyd. Growth dynamics of methanogens and sulfate reducers in natural marine sediments. Submitted

     Underwood, S., L. Lapham, A. Teske. K. G. Lloyd. 2016. Microbial community structure and methane-cycling activity of subsurface sediments at Mississippi Canyon 118 before the Deepwater Horizon disaster. Deep-Sea Research Part II. 129: 148-156. (pdf)

     Solden, L, K. G. Lloyd, K. Wrighton. 2016. The bright side of microbial dark matter: Lessons learned from the uncultivated majority. Current Opinion in Microbiology. 31:217-226. (pdf)

     Wasmund, K. C., M. Cooper, L. Schreiber, K. G. Lloyd, B. J. Baker, D. G. Petersen, B. B. Jorgensen, R. Stepanauskas, R. Reinhardt, A. Schramm, A. Loy, L. Adrian. 2016. Single-cell genome and group-specific dsrAB sequencing implicate marine members of the class Dehalococcoidia (Phylum Chloroflexi) in sulfur cycling. Mbio. 7:e00266-16. (pdf)

     Teske, A., D. de Beer, L. McKay, J. Biddle, M. Tivey, D. Hoer*, K. G. Lloyd, M. A. Lever, H. Røy, D. B. Albert, H. Mendlovitz, B. MacGregor. 2016. The Guaymas Basin hiking guide to hydrothermal mounds, chimneys and microbial mats: complex seafloor expressions of subsurface hydrothermal circulation. Frontiers in Microbiology. 7: 75. (pdf)

     Dowell, F.*, Z. Cardman*, Dasarathy, S., M. Y. Kellermann, J. S. Lipp, S. E. Ruff, J. F. Biddle, L. J. McKay, B. MacGregor, K. G. Lloyd, D. B. Albert, H. Mendlovitz, K. U. Hinrichs, A. Teske. 2016. Microbial communities in methane- and short chain alkane-rich hydrothermal sediments of Guaymas Basin. Frontiers in Microbiology. 7: 17. (pdf)

     Michalska, K., A. Steen, G. Chhor, M. Endres, J. Bird, K. G. Lloyd, A. Joachimiak. 2015. Discovery of novel aminopeptidase from “Microbial Dark Matter”, The FASEB Journal. 29:1-9. (pdf)

     Lever, M. A., K. Rogers, K. G. Lloyd, J. Overmann, B. Schink, R. K. Thauer, T. M. Hoehler, B. B. Jorgensen. 2015. The study of microbial life under extreme energy limitation: a synthesis of laboratory-based experiments and in-situ evidence from the deep biosphere. FEMS Microbiology Reviews. 39(5):688-728. (pdf)

     Wasmund, K., C. Algora, J. Muller, M. Krüger, K. G. Lloyd, R. Reinhardt, L. Adrian. 2015. Development and application of primers for the class Dehalococcoidetes (phylum Chloroflexi) enables deep insights into diversity and stratification of sub-groups in the marine subsurface. Environmental Microbiology17(10): 3540-3556. (pdf)

     Lloyd, K. G. 2014. Quantitative methods applied to methane-cycling archaea in the marine sediments of the White Oak River estuary. In Skovhus, T. L., Caffrey, S., and Hubert, C. Molecular Methods and Applications in Microbiology (Norfolk, UK: Caister Academic Press). 

     Orcutt, B. N., D. LaRowe, K. G. Lloyd, H. Mills, W. Orsi, B. Kiel-Reese, J. Sauvage. 2014. IODP Deep Biosphere Research workshop report – A synthesis of recent investigations, and discussion of new research questions and drilling targets. Scientific Drilling. 17: 61-66. 

     Lloyd, K. G. Importance of microbial quantification in the marine subseafloor. 2014. In Microbial Life of the Deep Biosphere, Jens Kallmeyer, Ed. (DeGruyter GmbH).

     Wasmund, K., L. Schreiber, K. G. Lloyd, D. G. Petersen, A. Schramm, R. Reinhardt, B. B. Jorgensen, L. Adrian. 2014. Genome sequencing of a single cell of the widely distributed marine subsurface Dehalococcoidetes, phylum Chloroflexi. ISME Journal. 8: 383-397. (pdf)

     Lloyd, K. G., L. Schreiber, D. G. Petersen, K. Kjeldsen, M. A. Lever, A. D. Steen, R. Stepanauskas, M. Richter, S. Kleindienst, S. Lenk, A. Schramm, B. B. Jørgensen. 2013. Predominant archaea in marine sediments degrade detrital proteins. Nature. 496: 215-219. (pdf)

     Lloyd, K. G., M. May, R. Kevorkian, A. D. Steen. 2013. A meta-analysis shows that archaea and bacteria have similar abundances in the subseafloor. Applied and Environmental Microbiology. 79: 7790-7799. (pdf)

     Meyer, S., G. Wegener, K. G. Lloyd, A. Teske, A. Boetius, A. Ramette. 2013. Microbial community connectivity across spatial scales and hydrothermal temperature gradients at Guaymas Basin. Frontiers in Marine Microbiology. 4: 207. (pdf)

     McKay, L. J., B. J. MacGregor, J. F. Biddle, D. B. Albert, H. P.Mendlovitz, D. R. Hoer, J. S. Lipp, K. G. Lloyd, A. P. Teske. 2012. Spatial heterogeneity and underlying geochemistry of phylogenetically diverse orange and white Beggiatoa mats in Guaymas Basin hydrothermal sediments. Deep-Sea Research Part I. 67:21-31. (pdf)

     Kubo, K., Lloyd, K. G., Biddle, J., Teske, A., Amann, R. Knittel, K. 2012. Archaea of the Miscellaneous Crenarchaeotal Group (MCG) are abundant, diverse, and widespread in marine sediments, ISME Journal. 6, 1949-1965. (pdf)

     Biddle, J. F., Z. Cardman, H. Mendlovitz, D. B. Albert, K. G. Lloyd, A. Boetius, A. Teske. 2012. Anaerobic oxidation of methane at different temperature regimes in Guaymas Basin hydrothermal sediments. ISME Journal. 6: 1018-1031. (pdf)

     Lloyd, K. G., M. J. Alperin, A. Teske. 2011. Environmental evidence for net methane production and oxidation in putative ANaerobic MEthanotrophic (ANME) archaea. Environmental Microbiology. 13(9): 2548-2564. (pdf)

     Lloyd, K. G., D. Albert, J. F. Biddle, J. Chanton, O. Pizarro, and A. Teske. 2010. Spatial structure and activity of sedimentary microbial communities underlying a Beggiatoa spp. mat in a Gulf of Mexico hydrocarbon seep. PLoS One, 5(1): e8738. (pdf)

     Lloyd, K. G., B. J. MacGregor, and A. Teske. 2010. Quantitative PCR methods for RNA and DNA in marine sediments: Maximizing yield while overcoming inhibition. FEMS Microbiology Ecology. 72: 143-151. (pdf)

     Joye, S. B., V. A. Samarkin, B. N. Orcutt, I. R. MacDonald, K. U. Hinrichs, M. Elvert, A. P. Teske, K. G. Lloyd, C. D. Meile, J. P. Montoya. 2009. Metabolic variability in seafloor brines revealed by carbon and sulphur dynamics. Nature Geoscience. 2(5): 349-354. (pdf)

     Edgcomb, V. P., S. J. Molyneaux, S. Böer, C. O. Wirsen, M. Saito, M. S. Atkins, K. G. Lloyd, A. Teske. 2007. Survival and growth of two heterotrophic hydrothermal vent archaea, Pyrococcus strain GB-D and Thermococcus fumicolans, under low pH and high sulfide concentrations in combination with high temperature and pressure regimes. Extremophiles. 11(2): 329-342. (pdf)

     Lloyd, K.G., L. Lapham, A. Teske. 2006. An anaerobic methane oxidizing community of ANME-1b archaea in hypersaline Gulf of Mexico sediments. Applied and Environmental Microbiology. 72: 7218-7230. (pdf)

     Biddle, J. F., J. S. Lipp, M. Lever, K. G. Lloyd, K. B. Sørensen, R. Anderson, H. F. Fredricks, M. Elvert, T. Kelly, D. P. Schrag, M. L. Sogin, J. Brenchley, A. Teske, C. H. House, K. U. Hinrichs. 2006. Heterotrophic archaea dominate sedimentary subsurface ecosystems off Peru. Proceedings of the National Academy of Sciences USA. 103(10): 3846-3851. (pdf)

     Lloyd, K.G., V.P. Edgcomb, S. M. Molyneaux, S. Böer, C.O. Wirsen, M. Atkins, A. Teske. 2005. Effect of dissolved sulfide, pH, and temperature on the growth and survival of marine hyperthermophilic archaea. Applied and Environmental Microbiology 71: 6383-6387. (pdf)

     Dhillon, A., M. Lever, K. Lloyd, D. Albert, M. L. Sogin, A. Teske. 2005. Methanogen diversity based on molecular characterization of Methyl Coenzyme M Reductase A (mcrA) genes in hydrothermal sediments of the Guaymas Basin. Applied and Environmental Microbiology. 71 (8): 4592-4601. (pdf)

     Edgcomb, V., S.J. Molyneaux, M.A. Saito, K. Lloyd, S. Böer, C.O. Wirsen, M.S. Atkins, and A. Teske. 2004. Sulfide ameliorates metal toxicity for deep-sea hydrothermal vent archaea. Applied and Environmental Microbiology 70:2551-2555. (pdf)

_____________________________________________________________________________________

University Teaching Experience

Research mentor and instructor, University of Tennessee                                                Sept 2012 – current

•  2 PhD students, 1 MS student, 10 BS students

•  Instructor for Micro 650: Topics in Microbial/Molecular Genetics, Micro 410: Microbial Physiology, Micro 310: Microbiology Survey, Biol 220: Intro Microbiology

Research mentor and guest lecturer, Aarhus University, Denmark                               Sept 2009 – Dec 2011

•  Guest lecturer for Microbial Diversity course

Research mentor, University of North Carolina                                                             Sept 2002 – Aug 2009

•  7 students from UNC-CH and 4 students from the University of Bremen, Germany

Guest lecturer, University of North Carolina, Chapel Hill, NC                                       Sept 2002 – Aug 2009

•  2 to 3 lectures per semester on microbial diversity, biogeochemistry, and oceanography

Writing Associate, Swarthmore College                                                                       Sept 1997 – May 2000

_____________________________________________________________________________________

Invited Lectures

Miami University, “Exploring the possible roles of uncultured marine sediment methanogens in reversible methanogenesis,” Oxford, OH. April 2015.

Argonne National Laboratory, “Exploring new branches of microbial life in the oceanic subsurface using single cell genomics,” Lemont, IL, October 2013

NASA Astrobiology Webinar, “How extensive, diverse and dynamic are subsurface and deep biospheres and how do they function?” Webcast in October 2013 archived at https://astrobiologyfuture.org/events/details/23

French Research Institute for Exploitation of the Sea (IFREMER), “ANMEs, methanogens, and other uncultured microbes,” Brest, France, December 2012

Woods Hole Oceanographic Institute public lecture, “Life in the deep ocean subseafloor: How Microbes make a living in some of the least hospitable environments on Earth,” Woods Hole, MA, July 2012

Woods Hole Oceanographic Institute scientific lecture, “Low energy life: Using transcriptomics and single cell genomics to study uncultured microbes in energy-limited systems,” Woods Hole, MA, July 2012

Oak Ridge National Laboratory, “Low energy life: Using single cell genomics and transcriptomics to study uncultured microbes in energy-limited systems”, Oak Ridge, TN, May 2012

Dark Energy Biosphere Institute Workshop, “Quantification of subsurface microorganisms,” University of North Carolina, Chapel Hill, NC, March 2011

North Carolina State University, “Determining the environmental functions of unknown organisms,” Dept. Marine, Earth and Atmospheric Sciences, Raleigh, NC, March 2011

University of Tennessee, “Determining the environmental functions of unknown organisms,” Knoxville, TN, January 2011

Skidaway Institute of Oceanography, “Important in nature but won’t grow in lab,” Savannah, GA, January 2011

University of Bremen, “Anaerobic methane oxidation,” Bremen, Germany, August 2005

Astrobiology Graduate Conference, “Investigating the potential for life in the hydrothermal vent subsurface,” Scripps Institute of Oceanography, San Diego, CA, August 2005

_____________________________________________________________________________________

Talks and Posters at Conferences and Workshops

2016

     Lloyd, K. G., J. Buongiorno, S. Turner, G. Webster, J. Bird, B. Reese, I. Marshall, M. Asai, A. Shumaker, T. Roy, A. Weightman, A. Schippers. Quantification of deep sedimentary microorganisms and single-cell genomic analysis from IODP Leg 347 Baltic Sea Paleoenvironment. Goldschmidt Conference. Yokohama, Japan. June 2016. Invited.

     Lloyd, K. G., TBD. ISME General Meeting. Montreal, Canada. August 2016. Invited.     

     Murray, P. N. Ferriero, J. Rosalsky, K. G. Lloyd, A. D. Steen. How do High School students respond to opportunities to collaborate with authentic scientific research in at-risk environments? Ocean Sciences Meeting. New Orleans, LA. February 2016.

     Buongiorno, J., S. Turner, G. Webster, A. Schumaker, T. Roy, A. Weightman, A. Schippers, K. G. Lloyd. Quantification of Deep Subseafloor Bacteria and Archaea – An Inter-Laboratory Comparison. Southeastern Biogeochemistry Symposium. Knoxville, TN 2016.

     Bird, J., B. Baker, M. Podar, K. G. Lloyd. Comparative Genomics Reveals Clues to the Evolutionary History of the Candidate Order Altiarchaeales. Southeastern Biogeochemistry Symposium. Knoxville, TN 2016.

2015

     Buongiorno, J., F. Martin, K. G. Lloyd. In situ DNA hybridized chain reaction (FISH-HCR) as a better method for bacterial and archaeal quantification within marine sediments. AGU. San Francisco, CA. December 2015. 

     Lloyd, K. G., R. Kevorkian, A. Shumaker, J. Bird. Unexpected microbial responses to redox shifts in marine sediments.  C-DEBI General Meeting, 10/4-10/7/2015, Marina, CA.

     Spirina, Elena, K. G. Lloyd, S. Pfiffner, E. Rivkina, T. Vishnivetskaya. Next generation research opportunities in microbiology and biogeosciences of Siberian permafrost. Permafrost in the XXI Century: Basic and Applied Research. Pushchino, Russia. September 2015.  

     Lloyd, K. G.  Introductory talk for Origin and Extent of Life. Kavli Institute’s 6th Indo-American Frontiers of Science Symposium. Irvine, CA. August 2015. Invited.

     Lloyd, K. G. J. Bird, S. Campagna, E. Tague, H. Castro, I. Marshall, B. Reese, G. Webster, A. Weightman. Preliminary results from meta-metabolomics and single cell genomics from the Baltic subsurface. IODP 2nd post-expedition symposium, Stockholm, Sweden. September 2015. Invited.

     Lloyd, K. G., J. Bird, S. Campagna, E. Tague, H. Castro, I. Marshall, B. Reese, G. Webster, A. Weightman. Abundance and functions of archaea and bacteria in the seabed. Microenergy Workshop, Sandbjerg, Denmark. September 2015. Invited.

     Lloyd, K. G. Using protein expression to explore novel functions in uncultured sediment microbes. C-DEBI Sediment Workshop. Los Angeles, CA. June 2015. Invited.

    Lloyd, K. G., J. Bird, I. Marshall, B. Reese, B. Baker, M. Podar. Obtaining single cell genomes from 85 meters below the seafloor, and using them to bin metagenomes. Single Cell Genomics Workshop, Boothbay Harbor, ME. June 2015.

     Lloyd, K. G. Linking single cell genomes of uncultured deep subsurface microbes to environmental functions using metagenomics. The Last frontier of Life ; Extremophiles - Expedition, Ecology, Adaptation, and Bioenergy production. Pyongchang, South Korea. May 2015. Invited.

     Lloyd, K. G. Linking deep subsurface microbial dark matter to their carbon cycling functions. Deep Carbon Observatory 2nd International Science Meeting, Munich, Germany. March 2015. Invited.

     Bird J. T., K. G. Lloyd, J. Buongiorno, H. F. Castro, E. D. Tague, S. R. Campagna, B. Reese, I. Marshall. Toward understanding the functional role of deeply-branching microbial communities in marine sediments from IODP Leg 347: Baltic Sea Paleoenvironment. Southeastern Biogeochemistry Symposium, Atlanta, GA. March 2015. 

     Buongiorno, J., A. Shumaker, J. Bird, K. Lloyd. Quantification of archaea and bacteria in deeply buried marine sediments from IODP Leg 347: Baltic Sea Paleoenvironment. Southeastern Biogeochemistry Symposium, Atlanta, GA. March 2015.

     Kevorkian, R. T., J. T. Bird*, K. G. Lloyd. Geochemical Control of Methanogenesis in Cape Lookout Bight, NC. Southeastern Biogeochemistry Symposium, Atlanta, GA. March 2015. 

     Keys, Jessica, T. Roy, K. G. Lloyd. Methanogenesis in marine sediment batch incubations. UT REU Symposium. August 2015. 

2014

     Lloyd, K. G., Carbon degradation mechanisms in uncultured marine organisms. American Society for Microbiology General Meeting, Boston, MA. May 2014. Invited.

     Lloyd, K. G., Methanogenesis in ANME archaea. C1 Compound Gordon Research Conference. Mt. Holyoke, MA. August 2014. Invited

     Lloyd, K.G., Exploring the genomes and novel enzyme functions of uncultured, deeply-branching archaea. International Society for Microbial Ecology. Seoul, South Korea. August 2014. Invited.

     Lloyd, K.G. Bird, J., Shumaker, A., IODP Shipboard Science Party Preliminary results from IODP Leg 347. IODP Workshop. Seoul, South Korea. August 2014. Invited.

     Lloyd, K.G., Exploring the possible roles of uncultured marine sediment methanogens in reversible methanogenesis. International Society for Subsurface Microbiology. Pacific Grove, CA. October 2014. Invited.

     Lloyd, K.G., Deep subsurface single cell genomes from IODP Leg 347: Baltic Sea Paleoenvironment  CDEBI Annual Meeting. Marina, CA. October 2014. Invited.

     Kevorkian, R. T., K. G. Lloyd, Identification of methanogens and controls on methane production in incubations of natural methane seep sediments. American Geophysical Union General Meeting, San Francisco, CA, December 2014. 

     Reese, B. K., L. Zinke, G. Carvalho, J. Amend, K. G. Lloyd, A. Shumaker, Ian Marshall, IODP Expedition 347 Shipboard Scientists, Active community structure analysis of deep subsurface sediments within Baltic Sea Basin. American Geophysical Union General Meeting, San Francisco, CA, December 2014. 

     Bird, J. T*, Expedition 347 Scientists, K. G. Lloyd, Genomic analysis of deeply-branching bacteria and archaea from IODP Leg 347: Baltic Sea Paleoenvironment. American Geophysical Union General Meeting, San Francisco, CA, December 2014. 

     Lloyd, K.G., J. Bird*, A. Shumaker, Expedition 347 Scientists. Potential physiologies of deep branches on the tree of life with deep subsurface samples from IODP Leg 347: Baltic Sea Paleoenvironment. American Geophysical Union General Meeting, San Francisco, CA, December 2014. Invited.

     Kevorkian, R. T., J. Bird, K. G. Lloyd. Geochemical control of methanogenesis in Cape Lookout Bight. Southeastern Biogeochemistry Symposium. April 2014.

     Bird, J., B. Baker, C. Lazar, M. Podar, A. Teske, K. G. Lloyd. Combining single cell and metagenomic sequencing: The first look at three members of the widely distributed, uncultured archaeal clade, SM1. Southeastern Biogeochemistry Symposium. April 2014.

     Papadoulis, S., J. Bird, K. G. Lloyd. Culturing the unculturable: growing archaea without the weeds. UT REU Symposium. August 2014. 

2013

     Steen, A. D., R. T. Kevorkian, E. Deas, K. G. Lloyd. Activities of extracellular peptidases in sediments of the White Oak River Estuary, NC. Center for Dark Energy Biosphere Investigations Annual Meeting. October 2013. Monterey, CA. 

    Steen, A.D, Kevorkian, R.T., Alperin, M.A., ,Lloyd, K.G.. Kinetics of extracellular peptidases in sediments of the White Oak River, NC, USA. American Geophysical Union General Meeting, December 2013, San Francisco, CA.

    Lloyd, K. G., J. Bird, L. Schreiber, D. G. Petersen, K. Kjeldsen, A. Schramm, R. Stepanauskas, B. B. Jørgensen. How do natural, uncultivated microbes interact with organic matter? Insights from single cell genomics and metagenomics. American Geophysical Union General Meeting, 12/3/13, San Francisco, CA

    Lloyd, K. G., M. May, A. D. Steen. A meta-analysis reveals methodological biases in quantitative methods for marine subsurface microbiology. Goldschmidt Conference. August 2013, Florence, Italy.

    Wasmund, K., C. Algora, M. Cookper, J. Mueller, M. Krueger, K. G. Lloyd, R. Reinhard, L. Adrian. Distributions and diversity of bacteria of the class Dehalococcoidia (phylum Chloroflexi) in the marine subsurface. Society for General and Applied Microbiology Germany Meeting, Bremen, Germany, March 2013

Meyer, S. G. Wegener, K. G. Lloyd, A. Teske, A. Boetius, A. Ramette. Microbial habitat connectivity in hydrothermal sediments of Guaymas Basin. SAME13 Aquatic Micrbogial Ecology. 

    Lloyd, K. G., M. May, A. D. Steen. A meta-analysis of quantitative microbiological methods in marine environments. American Society for Limnology and Oceanography Aquatic Sciences Meeting, New Orleans, LA, February 2013

2012

    Lloyd, K. G., Lars Schreiber, Dorthe Petersen, Andreas Schramm, Bo Barker Jørgensen. Potential environmental functions of widespread, abundant, uncultured marine archaea. American Geophysical Union General Meeting, San Francisco, CA, December 2012

    Lloyd, K. G. Are ANMEs methanogens? Gordon Research Seminar, Lewiston, ME, August, 2012 

    Lloyd, K. G., L. Schreiber, D. G. Petersen, K. Kjeldsen, M. A. Lever, A. D. Steen, R. Stepanauskas, M. Richter, S. Kleindienst, S. Lenk, A. Schramm, B. B. Jørgensen, Pinning down a metabolism for widespread, abundant, uncultured archaea through single cell genomic analysis. ISME Meeting, Copenhagen, Denmark, August 2012

    Petersen D.G., Lloyd K.G., Schreiber L., Richter M., Kjeldsen K., Lever M., Lenk S., Kleindienst S., Jørgensen B.B. and Schramm A. Phylogenetic identity and potential physiology linked in single microbial cells with focus on sulphate reducers. ISME Meeting, Copenhagen, Denmark, August 2012

    Schreiber L., Lloyd K.G., Petersen D.G., Richter M., Kjeldsen K., Lever M., Lenk S., Kleindienst S., Jørgensen B.B. and Schramm A. Reconstructuring genomeas of uncultured microorganisms from marine sediments using single cell genomics – an Aarhus Bay, Denmark, case study. ISME Meeting, Copenhagen, Denmark, August 2012

    Kjeldsen, Mark Lever, Karen Lloyd, Britta Gribsholt, Kristoffer Piil, Nils Risgaard-Petersen, Hans Røy, Waleed Al-Soud, Søren Sørensen, Bo Jørgensen. Changes in bacterial and archaeal community composition across geochemical zones in a coastal marine sediment. ISME Meeting, Copenhagen, Denmark, August 2012

    Wasmund K., Algora C., Schreiber L., Lloyd K. G., Petersen D., Andreas Schramm, Myriel Cooper, Richard Reinhardt, Bo Barker Jørgensen, Lorenz Adrian Insights into the ecological distribution and genomes of the enigmatic and widely distributed marine subsurface ‘Dehalococcoidetes’ (phylum Chloroflexi), ISME Meeting, Copenhagen, Denmark, August 2012

    Lloyd, K. G., et al., Single cell genomics of uncultured archaea. Goldschmidt Conference, Montreal, Canada, June 2012

    Alperin, M., T. Hoehler, K. G. Lloyd, On the reversibility of methane production from CO2 and H2: revisited. Microenergy 2012 Workshop, Aarhus, Denmark, May 2012

    Lloyd, K. G., Probing the genomic content of low-energy specialists. Microenergy 2012 Workshop, Aarhus, Denmark, May 2012

     McCully, E., J. Bird, K. G. Lloyd. Targeting cells of interest using fluorescently labeled peptides. UT REU Symposium. August 2015.

2011

    Petersen D.G., Schreiber L., Lloyd K., Schramm A. and Jørgensen B.B. Isolation of single microbial cells using a laser microdissection microscope (LMD). Nordic Environmental NUcleotide Network (NENUN), Reykjavik, Iceland, September 2011

    Lloyd, K. G., J. F. Biddle, K. Knittel, A. Teske. Environmental distribution, abundance, and activity of the Miscellaneous Crenarchaeotal Group. American Geophysical Union Fall Meeting. San Francisco, CA, December 2011

    Lloyd, K. G., M. Alperin, A. Teske. Transcriptional evidence for methane production in putative ANaerobic MEthanotrophic (ANME) archaea. Federation of European Microbiological Societies. 8th International Symposium of Subsurface Microbiology. Garmisch-Partenkirchen, Germany, September 2011

    Lloyd, K. G., M. Alperin, A. Teske. Transcriptional evidence for methane production in putative ANaerobic MEthanotrophic (ANME) archaea. Federation of European Microbiological Societies. FEMS Microbiology Congress. Geneva, Switzerland, June 2011

2010

    Lloyd, K. G., M. Alperin, A. Teske. Transcriptional evidence for methane production and methane oxidation in putative ANaerobic MEthanotrophic (ANME) archaea. American Geophysical Union Fall Meeting. San Francisco, CA, December 2010

2009

    Lloyd, K. G., D. Albert, J. Chanton, A. Teske. Spatial structure and activity of shallow subsurface microbial communities in a Gulf of Mexico methane seep. 19th Annual VM Goldschmidt Conference. Davos, Switzerland, June 2009

    Biddle, J. F., S. M. Huse, M. L. Sogin, K. G. Lloyd, A. Ramette, A. Boetius, A. Teske. Microbial consensus of marine methane seeps:  initial results. International Consensus of Marine Microbiology, February 2009

2008

    Lloyd, K. G., B. J. MacGregor, A. Teske. Quantification of archaeal RNA and DNA in marine sediments with qPCR and qRT-PCR. 12th ISME Meeting. Cairns, Australia, August 2008

    Lloyd, K. G., D. Albert, A. Teske. Seafloor bacterial mats as indicators of subsurface microbial activity in a Gulf of Mexico methane seep. ASLO/AGU Ocean Sciences. Orlando, FL, March 2008

2007

    Lloyd, K. G., A. Teske. Strategies for dealing with electron-acceptor limitation in estuarine sediments. International Workshop on Microbial Life under Extreme Energy Limitation. Aarhus, Denmark, October 2007 

2006

    Lloyd, K., A. Teske. Using functional gene transcripts to investigate anaerobic methane oxidation in estuarine sediments. 11th ISME Meeting, Vienna, Austria. August 2006

    Lloyd, K., A. Teske. Widespread active archaea in the deep and shallow subsurface. Astrobiology Science Conference, Washington, D.C., March 2006

2005

    Lloyd, K., L. Lapham, A. Teske. An Anaerobic Methane Oxidizing Community Dominated by ANME-1 in a Non-Hydrate-Associated Gulf of Mexico Sediment. NASA Astrobiology Institute General Meeting, Boulder, CO. April 2005

2003

    Lloyd, K., V. Edgcomb, S. Böer, S. Molyneaux, M. Atkins, C. Wirsen, A. Teske. Tolerances of hydrothermal vent archaea to deep subsurface conditions. NASA Astrobiology Institute General Meeting, Tempe, AZ. February 2003

2002

    Edgcomb V., S. Boer, S. Molyneaux, K. Lloyd, C. Wirsen, J. Erickson, M. Saito, M. Atkins, A. Teske. Hyperthermophiles of the hydrothermal vent subsurfaces: limits of life and extraterrestrial analogs. NASA Astrobiology Institute General Meeting, San Jose, CA. March 2002

    Edgcomb V., S. Boer, S. Molyneaux, K. Lloyd, C. Wirsen, J. Erickson, M. Saito, M. Atkins, A. Teske. Hyperthermophiles of the hydrothermal vent subsurface: limits of life. AGU/ASLO Ocean Sciences, Honolulu, HI. January 2002

2001

    Lloyd K., A. J. Engebretson, P. Adamczyk, C. Tomori, T. Lyons, C. L. Bennett. Out-of-pocket costs and lost wages associated with colorectal, breast, and prostate cancer care. American Society of Clinical Oncology, San Francisco, CA. March 2001

2000

    Lloyd K., K. P. Howard. Studying transmembrane helical peptides in magnetically oriented phospholipid bilayers. Biophysical Society General Meeting, New Orleans, LA. February 2000

_____________________________________________________________________________________

Field Experience

R/V Tyra, Aarhus Bay  (November 2010, March 2011)

R/V Susanne A Horsens, Aarhus Bay  (October 2009 & May 2010)

R/V Atlantis, DSV Alvin, East Pacific Rise and Guaymas Basin  (Nov-Dec 2008)

R/V Pelican, Gulf of Mexico MC118   (March 2006 & April 2008)

R/V Seward Johnson, DSV Johnson SeaLink, Gulf of Mexico MC118  (September 2006)

White Oak River Estuary, NC   (July 2005, Nov & Dec 2006, July 2008, Oct 2012)

_____________________________________________________________________________________

Outreach and Academic Service

Deep Carbon Observatory Executive Committee

Committee member: 7 PhD and 1 MS students (University of Tennessee), 1 MS student (Chesapeake Biological Laboratory), 1 PhD student (IFREMER, France)

Editor at mSystems

Editorial Board at Applied and Environmental Microbiology

Review manuscripts for: Antarctic Science, Applied and Environmental Microbiology, Astrobiology, BMC Microbiology, Geochimica et Cosmochimica Acta, Deep Sea Research, Environmental Science and Technology, FEMS Microbiology Ecology, FEMS Microbiology Letters, Frontiers in Microbiology, Geomicrobiology Journal, ISME Journal, Journal of Plankton Research, Life, Marine and Petroleum Geology, Microbial Ecology, Microbiome, mSystems, and Science

Review grants for: National Science Foundation, NASA Exobiology Program, L’Agence Nationale de la Recherche (France), NERC (UK)

Lead Organizer for: Southeastern Biogeochemistry Symposium, Knoxville, TN. Spring 2016

Co-Organizer for: C-DEBI Sediment Workshop, Los Angeles, CA. June 2015

Ethics Committee: Center for Dark Energy Biosphere Investigations

Theme leader for: Goldschmidt 2014, Sacremento, CA

Session chair for: ASLO Aquatic Sciences 2013, New Orleans, LA, ASLO Ocean Sciences 2014, Honolulu, HI, Goldschmidt 2014, Sacramento, CA, AGU 2016, San Francisco, CA

Collaborating with Malcom X. Shabazz High School on their Foundation for Newark’s Future project (2012-current)                                                                     

Volunteer for UT Darwin Day for education on evolution 2012 and 2015

Guest blogger, Extreme 2008 Deep-sea cruise

Developed and gave one-day workshops for elementary and middle school students:

“Diving in the Alvin submersible”, Montessori Farm School              (Spring 2009)

“Introduction to Oceanography” Culbreth Middle School       (April 2005 & 2007)

“Hydrothermal Vents and Alien Life” Stough Elementary School     (Spring 2003)

Science Judge for:

            University of Tennessee Undergraduate Science Competition, Eureca         (2013)

            Pathways Elementary School Science Fair                                           (April 2008)

National Ocean Sciences Bowl for high school students       (Spring 2006 & 2007) 

Graduate Student Action Group President                                                   (2006-2007)

Helped coordinate and participated in week-long scientific and cultural field trips to the NC coast and mountains, Culbreth Middle School (Spring 2004 & 2005)

Bi-weekly volunteer tutor for “Communities in Schools” Culbreth Middle School  (Fall 2002) 

© Karen Lloyd 2016